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| Pyroraptor Olympius. Only a few bones have been found of this species after a forrest fire in the Provence/South France. | |
NEWS :13. 10. 2005 - Sebastián Apesteguía, Federico Agnolín Novas and Peter Makovicky discover the remains of Buitreraptor gonzalezorum in the sandstones of La Buitrera, Rio Negro, in the north-west of Patagonia, Argentina. The new species displays characters also known from Troodontidae and Aves. It also is remarkable for it's completely unserrated teeth!2. 12. 2005 - A new specimen of Archaeopteryx has surfaced from a private collection and become available for research. Several characters, which were not well visible in previous known Archaeopteryx-Fossils or not well preserved, are clearly visible in the "new" excellently preserved specimen . As did the Deinonychosauria, Archaeopteryx possessed an enlarged hyperextendable second pedal ungual (claw on the 2nd toe) and the hallux (1st toe) was not opposable to the other toes. While the former is yet further proove that our modern birds are truly theropods, the latter points strongly towards Archaeopteryx never spent as much time in trees as many thought it would have (thus strengthening the "grounds up" - hypothesis concerning the evolution of powered flight)! |
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| The first step in order to understand the importance of the Deinonychosauria for palaeontology in general is to understand the relationships between the different species.
This cladogramm done by Michael Mortimer explains these relations very nicely (comments in brackets ( ) explain scientifical language): Format:
Alpha
|--Beta
`--Gamma
|--Delta
|*-Epsilon
`--Zeta
|?-Eta
`--Theta
Beta and Gamma are sister taxa (Species) within the clade (group) Alpha. Delta is within Gamma, but not a member of Zeta. Epsilon, however, is Gamma incertae sedis (uncertain position). Eta is only
tentatively placed within Zeta. Nomina nuda ("Naked name" species description considered invalid because either the specimen/s it was based on was/were lost or the material was considered inadequate) are non-italicized and placed in quotation marks, while unpublished taxa [as recognized by the ICZN] are designated
the same way, but followed by an asterisk. Nomina dubia (Species "in doubt" meaning it's not for certain, wether this actually is a valid species) are not distinguished from valid taxa, as which taxa are indeterminate is a subjective question.
Theropoda | Avepoda Averostra Neotheropoda | Ceratosauroidea | Tetanurae | Avetheropoda | Coelurosauria | Avipluma Tyrannoraptora | Metatheropoda | Maniraptoriformes | Maniraptora | Paraves Eumaniraptora | |--Deinonychosauria | Dromaeosauridae | |--Microraptoria | | |--Graciliraptor lujiatunensis | | |--Sinornithosaurus millenii | | `--+--Cryptovolans pauli | | |--Microraptor gui | | `--Microraptor zhaoianus | |--Bambiraptor feinbergi | |*?"Airakoraptor" | |*-"Dromaeosaurus" gracilis | |*?"Ichabodcraniosaurus" | |*-Itemirus medullaris | |*-"Laelaps" explanatus | `--+--Velociraptorinae | | `--Velociraptor mongoliensis | |?-Saurornitholestes langstoni | |*-"Koreanosaurus koreanensis" | |*-Ornithodesmus cluniculus | |*-Pyroraptor olympius | |*-Variraptor mechinorum | `--Dromaeosaurinae | |--Deinonychus antirrhopus | `--+--Atrociraptor marshalli | |?-Dromaeosauroides bornholmensis | `--+--Dromaeosaurus albertensis | |?-Adasaurus mongoliensis | |--Achillobator giganticus | `--Utahraptor ostrommaysorum |--Troodontidae | |*-Yaverlandia bitholus | |--Mei long | `--+--Sinovenator changiae | |--Jinfengopteryx elegans | `--+--Sinusonasus magnodens | |--Koparion douglassi | |*?"Ornithomimus" tenuis | `--+--Byronosaurus jaffei | `--+--Sinornithoides youngi | `--+*-Borogovia gracilicrus | |*-Saurornithoides? asiamericanus | |*-Saurornithoides? isfarensis | |*-Tochisaurus nemegtensis | |--Troodon formosus | `--+--Saurornithoides mongoliensis | `--Saurornithoides junior `--Avialae
The main informations we can get out of this cladogramm are the following:
Distribution: Fossil remains of Deinonychosauria (including Troodontidae) have been described from all over the world.
Timeline: The following table shows when the different species lived and who described them ("correctly" - according to present knowledge) for the first time.
| |
| Species name | Author/s | Described in | Location | Time period |
| Troodontidae | ||||
|---|---|---|---|---|
| Sinovenator changii | Xu, Norell, Wang, Makovicky & Wu | 2002 | Lujiatun Beds of Yixian Formation, Liaoning, China | Hauteverian, Early Cretaceous |
| Koparion douglassi | Chure | 1994 | Brushy Basin Member of the Morrison Formation, Utah, U.S. | Tithonian, Late Jurassic |
| Saurornithoides asiamericanus | Nessov / Olshevsky | 1985 / 1991 | Khodzhakul Formation, Sheykhdzeyli, Uzbekistan | Early Cenomanian, Late Cretaceous |
| Saurornithoides isfarensis | Nessov / Olshevsky | 1995 / 2000 | Yalovach Formation, Kyzyl-pilyal, Tadhzikistan | Early Santonian, Late Cretaceous |
| Sinornithoides youngi | Russell & Dong | 1994 | Ejinhoro Formation, Inner Mongolia; Xinminbao Group, Gansu, China | Albian, Early Cretaceous |
| Borogovia gracilicrus | Osmolska | 1987 | Nemegt Formation, Mongolia | Late Campanian - Early Maastrichtian, Late Cretaceous |
| Tochisaurus nemegtensis | Kurzanov & Osmolska | 1991 | Nemegt Formation, Mongolia | Late Campanian - Early Maastrichtian, Late Cretaceous |
| Troodon formosus | Leidy | 1856 | Dinosaur Park Formation, Oldman Formation, Two Medicine Formation, Alberta, Canada; Prince Creek Formation, Alaska, Judith River Group, Upper Two Medicine Formation, Montana, Fruitland Formation, Kirtland Formation, Utah, U.S. | Late Campanian - Early Maastrichtian, Late Cretaceous |
| Ornithomimus tenius | Marsh | 1890 | Judith River Formation, Montana, U.S. | Late Campanian, Late Cretaceous |
| Jinfengopteryx elegans | Qiang, Shu'an, Junchang, Hailu, Wen, Yongqing & Yanxue | 2005 | Qiaotu Formation, Hebei, China | Late Jurassic - Early Cretaceous |
| Byronosaurus jaffei | Norell, Makovicky & Clark | 2000 | Ukhaa Tolgod, Mongolia | Late Cretaceous |
| Mei long | Xing Xu & Mark A. Norell | 2004 | Lujiatun, western Liaoning, China | Berreasian - Hauteverain, Early Cretaceous |
| Microraptoria | ||||
| Microraptor zhaoianus | Xu, Zhou & Wang | 2000 | Yixian Formation, Liaoning, China | Barremian, Early Cretaceous |
| Microraptor gui | Xu, Zhou, Wang, Kuang, Zhang & Du | 2003 | Jiufotang Formation, Liaoning, China | Early Aptian, Early Cretaceous |
| Cryptovolans pauli | Czerkas, Zhang, Li & Li | 2002 | Jiufotang Formation, Liaoning, China | Early Aptian, Early Cretaceous |
| Sinornithosaurus Millenii | Xu, Wang & Wu | 1999 | Jianshangu Member of Yixian Formation, Liaoning, China | Middle Barremian, Early Cretaceous |
| Graciliraptor lujiatonensis | Xu & Wang | 2004 | Yixian Formation, Liaoning, China | Hauterivian, Early Cretaceous |
| Dromaeosauridae | ||||
| Bambiraptor feinbergi | Burnham, Derstler, Currie, Bakker, Zhou & Ostrom | 2000 | Upper Two Medicine Formation, Montana, U.S. | Campanian, Late Cretaceous |
| Variraptor mechinorum | Le Loeff & Buffetaut | 1998 | Gres a Reptiles Formation, France | Late Campanian - Early Maastrichtian, Late Cretaceous |
| Pyroraptor olympius | Allain & Taquet | 2000 | La Boucharde, Bouches-du-Rhone, France | Late Campanian - Early Maastrichtian, Late Cretaceous |
| Ornithodesmus cluniculus | Seeley | 1887 | Upper Weald Clay, England | Barremian, Early Cretaceous |
| Dromaeosaurus albertensis | Matthew & Brown | 1922 | Dinosaur Park Formation, Foremost Formation, Oldman Formation, Alberta, Canada; Prince Creek Formation, Alaska, Judith River Group, Lower Two Medicine Formation, Montana, U.S. | Late Campanian - Maastrichtian, Late Cretaceous |
| Dromaeosauroides bornholmensis | Christiansen & Bonde | 2003 | Jydegaard Formation, Denmark | Late Berriasian, Early Cretaceous |
| Deinonychus antirrhopus | Ostrom | 1969 | Unknown Formation, Arizona, Arundel Formation, Maryland, Cloverly Formation, Montana, Antlers Formation, Oklahoma, Trinity Group, Texas, middle Cedar Mountain Formation, Utah, U.S. | Aptian - Albian, Early Cretaceous |
| Saurornitholestes langstoni | Sues | 1978 | Dinosaur Park Formation, Foremost Formation, Milk River Formation, Oldman Formation, Two Medicine Formation, Alberta, Canada; Prince Creek Formation, Alaska, Judith River Group, Lower Two Medicine Formation, Montana, Menefee Formation, New Mexico, U.S. | Early Campanian - Maastrichtian, Late Cretaceous |
| Velociraptor mongoliensis | Osborn | 1924 | Barun Goyot Formation, Djadochta Formation, Mongolia | Late Santonian - Early Campanian, Late Cretaceous |
| Ichabodcraniosaurus | Novacek | 1996 | Djadochta Formation, Mongolia | Campanian, Late Cretaceous |
| Airakoraptor | Perle, Norell & Clark | 1999 | Mongolia | Late Cretaceous |
| Adasaurus mongoliensis | Barsbold | 1983 | Nemegt Formation, Mongolia | Middle Maastrichtian, Late Cretaceous |
| Achillobator giganticus | Perle, Norell & Clark | 1999 | Bayan Shireh Formation, Mongolia | Santonian - Campanian, Late Cretaceous |
| Utahraptor ostrommaysorum | Kirkland, Gaston & Burge | 1993 | Yellow Cat Member of Cedar Mountain Formation, Utah, U.S. | Barremian, Early Cretaceous |
| Neuquenraptor argentinus | Novas & Pol | 2005 | Portezuelo Formation, Neuquen Province, Argentina | Coniacian, Early Cretaceous |
| Unnamed dromaeosaur* | Rauhut & Werner | 1995 | Wadi Milk Formation, Sudan | Cenomanian, Late Cretaceous |
| * = this specimen was added in order to illustrate the presence of dromaeosaurids on the African continent. | ||||
| Besides the fossil material that was used to describe and name the species listed above, there still exists plenty of undescribed and unnamed
fossil material. Some of these fossils belong to the Troodontidae and Dromaeosauridae and mostly consist of teeth. The time periods these unnamed and undescribed fossils belong to range from
the late Jurassic to the late Cretaceous time. All this data so far available allows to conclude that Deinonychosauria were present from the late Jurassic until the end of the Cretaceous time.
This means they were around for an impressive 85 million years which underlines the success of the group as such! The following table gives a good oversight throughout what timespan Deinonychosauria and Troodontidae were actually present on our planet. It covers only those periods during which Deinonychosauria or Troodontidae were present. The Mesozoic era covers also the Triassic period and those parts of the Jurassic which are actually not displayed in this table. The Titonian age began about 150,7 million years ago. |
| The period of time during which the Deinonychosauria were present on earth. Age estimates are in millions of years. | |||||
|---|---|---|---|---|---|
| Era | Period | Epoch | Age | End | |
| Mesozoic | Cretaceous | Late | Senonian | Maastrichtian | 65 |
| Campanian | 71 | ||||
| Santonian | 83,5 | ||||
| Coniacian | 85,8 | ||||
| Gallic | Turonian | 89,9 | |||
| Cenomanian | 93,5 | ||||
| Early | Albian | 98,9 | |||
| Aptian | 112,2 | ||||
| Barremian | 121 | ||||
| Neocomian | Hauterivian | 127 | |||
| Valanginian | 132 | ||||
| Berriasian | 137 | ||||
| Jurassic | Late | Malm | Titonian | 144,2 | |